It is a convenient reference point with which to measure many voltages in the circuit. It may or may not also be connected to the Earth with a grounding wire.
Once you pick a convention (electrons travel around the circuit one way, holes and 'conventional current' the other way) just be consistent it its application. If you want to think of current as a flow of electrons, then current will take all paths that connect a more negative voltage to a more positive one. Ground, by definition, is a zero volt reference point.
You have to separate in your mind voltages from currents. Nodes can be have a more positive or negative voltage than ground. Pick any two nodes, and one may be the more positive and the other the more negative, or vice versa. This potential difference is what drives current through the things connected between those nodes. Currents do not have the same sort of positive and negative polarities that voltages do. Current polarities just refer to whether an actual current goes the assumed way (positive current) or the opposite way (negative current) so that you can add them up correctly for things like Kirchoff's law (the sum of all currents into a node is zero).
Have I helped or added to your confusion?